/* * ContinuousBranchRates.java * * Copyright (C) 2002-2009 Alexei Drummond and Andrew Rambaut * * This file is part of BEAST. * See the NOTICE file distributed with this work for additional * information regarding copyright ownership and licensing. * * BEAST is free software; you can redistribute it and/or modify * it under the terms of the GNU Lesser General Public License as * published by the Free Software Foundation; either version 2 * of the License, or (at your option) any later version. * * BEAST is distributed in the hope that it will be useful, * but WITHOUT ANY WARRANTY; without even the implied warranty of * MERCHANTABILITY or FITNESS FOR A PARTICULAR PURPOSE. See the * GNU Lesser General Public License for more details. * * You should have received a copy of the GNU Lesser General Public * License along with BEAST; if not, write to the * Free Software Foundation, Inc., 51 Franklin St, Fifth Floor, * Boston, MA 02110-1301 USA */ package dr.evomodel.branchratemodel; import dr.evolution.tree.NodeRef; import dr.evolution.tree.SimpleTree; import dr.evolution.tree.Tree; import dr.evomodel.tree.TreeModel; import dr.evomodel.tree.TreeParameterModel; import dr.evomodelxml.branchratemodel.ContinuousBranchRatesParser; import dr.inference.distribution.ParametricDistributionModel; import dr.inference.model.*; /** * * Uses the implementation of continuous branch rates described in: * Li, W.L.S. and A.J. Drummond, Model Averaging and Bayes Factor * Calculation of Relaxed Molecular Clocks in Bayesian Phylogenetics. * Molecular Biology and Evolution, 2012. 29(2): p. 751-761. * * @author Wai Lok Sibon Li * @version $Id: ContinuousBranchRates.java,v 1.11 2006/01/09 17:44:30 rambaut Exp $ */ public class ContinuousBranchRates extends AbstractBranchRateModel { private final ParametricDistributionModel distributionModel; // The rate categories of each branch //d final TreeParameterModel rateCategories; final TreeParameterModel rateCategoryQuantiles; //private final int categoryCount; //private final double step; private final double[] rates; private boolean normalize = false; private double normalizeBranchRateTo = Double.NaN; private double scaleFactor = 1.0; private TreeModel treeModel; private Tree tree; //overSampling control the number of effective categories public ContinuousBranchRates( TreeModel tree, /*Parameter rateCategoryParameter, */ Parameter rateCategoryQuantilesParameter, ParametricDistributionModel model /*int overSampling*/) { this(tree, /* rateCategoryParameter, */rateCategoryQuantilesParameter, model, /*overSampling, */false, Double.NaN); } public ContinuousBranchRates( TreeModel tree, /* Parameter rateCategoryParameter, */ Parameter rateCategoryQuantilesParameter, ParametricDistributionModel model, /*int overSampling,*/ boolean normalize, double normalizeBranchRateTo) { super(ContinuousBranchRatesParser.CONTINUOUS_BRANCH_RATES); //d this.rateCategories = new TreeParameterModel(tree, rateCategoryParameter, false); this.rateCategoryQuantiles = new TreeParameterModel(tree, rateCategoryQuantilesParameter, false); rates = new double[tree.getNodeCount()]; this.normalize = normalize; this.treeModel = tree; this.distributionModel = model; this.normalizeBranchRateTo = normalizeBranchRateTo; this.tree = new SimpleTree(tree); //Force the boundaries of rateCategoryParameter to match the category count //d Parameter.DefaultBounds bound = new Parameter.DefaultBounds(categoryCount - 1, 0, rateCategoryParameter.getDimension()); //d rateCategoryParameter.addBounds(bound); //rateCategoryQuantilesParameter.; Parameter.DefaultBounds bound = new Parameter.DefaultBounds(1.0, 0.0, rateCategoryQuantilesParameter.getDimension()); rateCategoryQuantilesParameter.addBounds(bound); /*if(rateCategoryQuantilesParameter.getBounds()==null) { System.out.println("oh NO!!! " + rateCategoryQuantilesParameter.getBounds().getLowerLimit(0) + "\t" + rateCategoryQuantilesParameter.getBounds().getUpperLimit(1)); }*/ /* for (int i = 0; i < rateCategoryParameter.getDimension(); i++) { int index = (int) Math.floor((i + 0.5) * overSampling); rateCategoryParameter.setParameterValue(i, index); } */ addModel(model); // AR - commented out: changes to the tree are handled by model changed events fired by rateCategories // addModel(tree); //d addModel(rateCategories); addModel(rateCategoryQuantiles); //addModel(treeModel); // Maybe // AR - commented out: changes to rateCategoryParameter are handled by model changed events fired by rateCategories // addVariable(rateCategoryParameter); if (normalize) { tree.addModelListener(new ModelListener() { public void modelChangedEvent(Model model, Object object, int index) { computeFactor(); } public void modelRestored(Model model) { computeFactor(); } }); } setupRates(); } // compute scale factor private void computeFactor() { //scale mean rate to 1.0 or separate parameter double treeRate = 0.0; double treeTime = 0.0; //normalizeBranchRateTo = 1.0; for (int i = 0; i < treeModel.getNodeCount(); i++) { NodeRef node = treeModel.getNode(i); if (!treeModel.isRoot(node)) { //d int rateCategory = (int) Math.round(rateCategories.getNodeValue(treeModel, node)); //d treeRate += rates[rateCategory] * treeModel.getBranchLength(node); treeTime += treeModel.getBranchLength(node); // d System.out.println("rates and time\t" + rates[rateCategory] + "\t" + treeModel.getBranchLength(node)); } } //treeRate /= treeTime; scaleFactor = normalizeBranchRateTo / (treeRate / treeTime); System.out.println("scaleFactor\t\t\t\t\t" + scaleFactor); } public void handleModelChangedEvent(Model model, Object object, int index) { if (model == distributionModel) { setupRates(); fireModelChanged(); } //else if (model == rateCategories) { // AR - commented out: if just the rate categories have changed the rates will be the same // setupRates(); // fireModelChanged(null, index); //} else if (model == rateCategoryQuantiles) { setupRates(); // Maybe //rateCategories.fireModelChanged(); fireModelChanged(null, index); } /*else if (model == treeModel) { setupRates(); // Maybe }*/ } protected final void handleVariableChangedEvent(Variable variable, int index, Parameter.ChangeType type) { // AR - commented out: changes to rateCategoryParameter are handled by model changed events //setupRates(); // Maybe } protected void storeState() { //setupRates(); // Maybe } protected void restoreState() { setupRates(); } protected void acceptState() { //setupRates(); // Maybe } public double getBranchRate(final Tree tree, final NodeRef node) { assert !tree.isRoot(node) : "root node doesn't have a rate!"; //int rateCategory = (int) Math.round(rateCategories.getNodeValue(tree, node)); //System.out.println("dslkjafsdf " + node.getNumber()); return rates[node.getNumber()] * scaleFactor; } /** * Calculates the actual rates corresponding to the category indices. */ protected void setupRates() { //rateCategoryQuantiles. //double z = step / 2.0; for (int i = 0; i < tree.getNodeCount(); i++) { //rates[i] = distributionModel.quantile(rateCategoryQuantiles.getNodeValue(rateCategoryQuantiles.getTreeModel(), rateCategoryQuantiles.getTreeModel().getNode(i) )); if (!tree.isRoot(tree.getNode(i))) { //System.out.println(rateCategoryQuantiles.getNodeValue(tree, tree.getNode(i))); rates[tree.getNode(i).getNumber()] = distributionModel.quantile(rateCategoryQuantiles.getNodeValue(tree, tree.getNode(i))); } // System.out.println("road " + i + "\t" + rateCategoryQuantiles.getNodeValue(tree, tree.getNode(i) ) + "\t" + tree.toString() + "\t" + tree.toString()); //System.out.println("road " + i + "\t" + rateCategoryQuantiles.getNodeValue(rateCategoryQuantiles.getTreeModel(), rateCategoryQuantiles.getTreeModel().getNode(i) ) + "\t" + rateCategoryQuantiles.getTreeModel().toString() + "\t" + treeModel.toString()); //System.out.print(rates[i]+"\t"); //z += step; //rates[i] = distributionModel.quantile(z); //System.out.println("ape " + tree); } //System.out.println("\n"); //System.out.println(); if (normalize) computeFactor(); } }